Guide to Liverworts of Oregon: Cephaloziella divaricata (Sm.) Schiffn.
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Synonym: Cephaloziella scabra, Cephaloziella byssacaea
Special Status: None.
Recognition: Deeply bilobed leaves, lobes triangular, underleaves consistently present on sterile shoots. Dioicous. Leaves often with abaxial papillae or projections.
Distribution: On soil, wood, charcoal, and rock; at all elevations.
Comments: One of the most widespread of liverworts, both geographically and ecologically. Prior to 2017 it had been my practice to name all sterile material that fits this description as Cephaloziella divaricata even though it could theoretically be Cephaloziella stellulifera. The treatment here has been updated to incorporate C. stellulifera.
It is critical to recognize the paroicous inflorescences of Cephaloziella stellulifera as distinct from Cephaloziella divaricata. Cephaloziella divaricata is always described as dioicous but occasional autoicous plants have been observed. In either case, the presence of abundant, solitary androecia in a collection will rule out the paroicous species. Cephaloziella stellulifera is distinguished by edentate female bracts and long perianth mouth cells which may be free at the tip. Its leaves are consistently smooth abaxially in contrast to the forms of Cephaloziella divaricata with armed leaves. Female bracts of Cephaloziella divaricata are always dentate and the perianth mouth cells are mostly short and form a crenulate margin without free tips.
For an overview of full resolution images, consult the ancillary file: Cephaloziella image atlas.
New River, Coos Co., Oregon. DHW
m0783a.
Malakof Diggings, Nevada County, California. C.Beyer 7051.
Cephaloziella divaricata. Boulder County, Colorado. Weber B-48266 (ALA).
Cephaloziella divaricata -- Stink Creek, Grant County, Oregon. D.Stone 8403-3.
Cephaloziella divaricata -- Stink Creek, Grant County, Oregon. D.Stone 8403-3.
Cephaloziella divaricata -- Stink Creek, Grant County, Oregon. D.Stone 8403-3.
Cephaloziella divaricata -- Oil bodies in leaf. Stink Creek, Grant County, Oregon. D.Stone 8403-3.
Cephaloziella divaricata -- Heavily gemmiferous colony. Stink Creek, Grant County, Oregon. D.Stone 8403-3.
Cephaloziella divaricata -- Stink Creek, Grant County, Oregon. D.Stone 8403-3.
Cephaloziella divaricata -- Stink Creek, Grant County, Oregon. D.Stone 8403-3.
Cephaloziella divaricata Gynoecium, stained with toluidine blue. Hunter Creek, Curry County, Oregon. DHW 1944.
Cephaloziella divaricata Perianth mouth, stained with toluidine blue. Hunter Creek, Curry County, Oregon. DHW 1944.
Cephaloziella divaricata Shoots 10X, stained with toluidine blue. Hunter Creek, Curry County, Oregon. DHW 1944.
Cephaloziella divaricata Leaf 40X, stained with toluidine blue. Hunter Creek, Curry County, Oregon. DHW 1944.
Cephaloziella divaricata Shoot with perianth. Note absence of gibbous bractes below perianth. Hunter Creek, Curry County, Oregon. DHW 1966.
Cephaloziella divaricata Shoot with perianth. Note absence of gibbous bractes below perianth. Hunter Creek, Curry County, Oregon. DHW 1966.
Cephaloziella divaricata Shoot 40X. Sleeping Beauty Trail, Skamania County, Washington. Norris 104455 (UC).
Cephaloziella divaricata Lateral aspect. Sleeping Beauty Trail, Skamania County, Washington. Norris 104455 (UC).
Cephaloziella divaricata Leaf, extreme development of branched projections. Sleeping Beauty Trail, Skamania County, Washington. Norris 104455 (UC).
Cephaloziella divaricata. Small shoot. Andrews Experimental Forest, Lane County, Oregon. Jonsson 201A/94.
Cephaloziella divaricata. Small shoot. Andrews Experimental Forest, Lane County, Oregon. Jonsson 201A/94.
Cephaloziella divaricata. Robust shoot with leaf cells measured on three leaves (measurement in faint pink on each cell measured). Andrews Experimental Forest, Lane County, Oregon. Jonsson 201A/94.